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Miss Chen
2018年06月18日
Miss Chen
Description: This perennial plant is 3-5' tall and unbranched, except for flowerhead-bearing stalks near the apex of the plant. The central stem is stout, terete, and either purple-spotted or purple; it is often covered with short fine hairs. At intervals along the central stem, there are whorls of 4-5 leaves that are yellowish green or green. These leaves are up to 7" long and 2½" across; they are lanceolate to broadly elliptic and serrated along their margins. The upper leaf surfaces have conspicuous venation. The petioles of the leaves are short (less than ¼" in length). The central stem terminates in a rather flat-headed panicle of flowerheads spanning 3-6" across. Small clusters of flowerheads may occur below on separate stalks. Each narrow flowerhead is about 1/3" (8 mm.) long, consisting of 8-20 pink to purplish pink disk florets and no ray florets. Each tiny disk floret has a narrowly tubular corolla with 5 spreading lobes along its upper rim; exerted from the corolla, there is a strongly exerted style that is divided into filiform parts. The narrow bracts (phyllaries) at the base of each flowerhead are pink or purplish pink, like the disk florets. The flowering stalks are purple-spotted to purple and they are often covered with short fine hairs. The blooming period occurs from mid- to late summer, lasting about 3-4 weeks. The flowerheads are often fragrant. The florets are replaced by achenes with small tufts of bristly hair; they are dispersed by the wind. The root system is fibrous and rhizomatous. This plant often forms small clonal colonies. Cultivation: The preference is full or partial sun, wet to moist conditions, and a mineral-rich soil containing silty or sandy loam. Spotted Joe-Pye Weed is more tolerant of water-logged conditions than most plants. Range & Habitat: The native Spotted Joe-Pye Weed occurs occasionally in the northern half of Illinois, while in the southern half of the state it is rare or absent (see Distribution Map). Habitats include wet black soil prairies, wet sand prairies, sedge meadows, marshes, fens, and swampy thickets with small trees or shrubs. Spotted Joe-Pye Weed is partial to sandy wetlands, but it is also found in non-sandy wetlands. It is usually found in high quality natural areas, rather than degraded habitats with a history of disturbance.
Faunal Associations: The nectar of the flowers attracts honey bees, bumblebees, long-horned bees (Melissodes spp.), leaf-cutting bees (Megachile spp.), bee flies, butterflies, skippers, and moths. Some bees may also collect pollen. The following leaf beetles have been observed to feed on Spotted Joe-Pye Weed: Exema dispar, Ophraella notata, and Sumitrosis inaequalis (Clark et al., 2004). An uncommon aphid, Aphis vernoniae, sucks the plant juices. The caterpillars of some moth species feed on various parts of Eutrochium spp. (Joe-Pye Weed species). They include Schinia trifascia (Three-Lined Flower Moth; feeds on florets), Carmenta bassiformis (Eupatorium Borer Moth; bores through roots), Phragmatobia fuliginosa (Ruby Tiger Moth; feeds on foliage), and Eupithecia miserulata (Common Pug; feeds on florets). The seeds of Joe-Pye Weed species are a minor source of food to the Swamp Sparrow. The foliage is not preferred as a food source for mammalian herbivores, although it may be browsed upon occasion by deer, rabbits, or livestock. Photographic Location: Cowe's Bog at the Indiana Dunes National Lakeshore in NW Indiana. Cowe's Bog is a sandy fen.
Comments: Recently, species of Joe-Pye Weed have been separated from the Bonesets (Eupatorium spp.); they have been re-assigned to the genus Eutrochium. Some sources of information still refer to this species as Eupatorium maculatum, and sometimes it is also referred to as Eupatoriadelphus maculatus. In Illinois, species of Joe-Pye Weed differ from the Bonesets by their whorled leaves, while the latter group of plants usually have opposite leaves. Spotted Joe-Pye Weed (Eutrochium maculatum) differs from Hollow-stemmed Joe-Pye Weed (Eutrochium fistulosum) and Sweet Joe-Pye Weed (Eutrochium purpureum) by its rather flat-headed panicles of flowerheads, purple-spotted or purplish stems that are often pubescent, and flowerheads consisting of 8-20 disk florets. Other species of Joe-Pye Weed within the state have more dome-shaped panicles of flowerheads, stems that are hairless, and flowerheads consisting of 4-7 disk florets. Spotted Joe-Pye Weed also tends to be shorter.
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Miss Chen
2018年06月18日
Miss Chen
Description: This perennial wildflower is 3-9' tall and largely unbranched, except for some lateral ascending stems along the upper one-third of its length. The central stem is dark purple to pale purplish white, stout, terete, glabrous, and glaucous; the interior of the central stem is mostly hollow. The lateral stems are similar, except they are more narrow. Whorls of 4-7 leaves (usually 5-6) occur at intervals along the central stem; whorls of smaller leaves also occur along some of the lateral stems. Individual leaves are up to 9" long and 3" across; they are elliptic in shape and crenate-serrate along their margins. The upper leaf surface is medium green and glabrous; it is creased along the veins. The lower leaf surface is pale green and mostly glabrous, except for hairs along the lower sides of the veins. The petioles are up to ½" long, pale purple to dark purple, and stout. The central stem terminates in a panicle of flowerheads up to 1½' long and 1' across; the upper lateral stems often terminate in panicles of flowerheads as well, except they are smaller in size. Individual panicles are rather open and dome-shaped at the top; they vary in color from pale pink-lavender to deep rosy pink. Individual flowerheads are about 9 mm. (1/3") long and 3 mm. (1/8") across, consisting of 5-7 disk florets and no ray florets. Each disk floret consists of a narrowly cylindrical corolla with 5 upright lobes, 5 inserted stamens, and a pistil with an exserted bifurcated style. The corolla is pink-lavender to rosy pink. At the base of each flowerhead, there are overlapping bracts (phyllaries) that are appressed together in several series. These bracts are usually linear-oblong, white with pink tints to light pink, and glabrous. The branches of each panicle are widely spreading to ascending; they are usually dark purple and finely pubescent. The blooming period occurs from mid-summer to early fall and lasts about 1 month. The flowerheads are often mildly fragrant. Afterwards, the disk florets are replaced by small bullet-shaped achenes (about 3 mm. long) with tufts of bristly hair. They are distributed by the wind. The root system is mostly fibrous, although sometimes rhizomes are produced.Distribution Map Cultivation: The preference is full or partial sun and wet to moist conditions; the soil can contain loam, sand, or gravel. There is considerable variation in the size of individual plants. Exposure to high winds can cause some plants to topple over. Standing water is tolerated if it is temporary. Range & Habitat: The native Hollow-Stemmed Joe-Pye Weed is an uncommon plant that is found in the southern tip of Illinois, SE Illinois, and widely scattered areas elsewhere. Habitats include wet sand prairies, seeps and springs in partially wooded areas, soggy thickets, fens, rocky sandstone ravines along streams, and low areas along railroads and roadsides. Sometimes this wildflower is cultivated in gardens, although many cultivars display some evidence of hybridization. Faunal Associations: The flowerheads are visited by honeybees, bumblebees, and other long-tongued bees; other floral visitors include bee flies (Bombyliidae), butterflies, skippers, and moths. Most of these visitors obtain nectar from the flowerheads, although some bees also collect pollen. A modest number of insects feed on the foliage, flowers, stems, or roots of Hollow Joe-Pye Weed and other members of this genus. They include caterpillars of the following moths: Carmenta bassiformis (Eupatorium Borer Moth), Condica vecors (Dusky Groundling), Schinia trifascia (Three-Lined Flower Moth), Papaipema eupatorii (Joe-Pye Weed Borer), Perigea xanthioides (Red Groundling), and Phragmatobia fuliginosa (Ruby Tiger Moth). Other insect feeders include the aphid Aphis vernoniae, the treehopper Entylia bactriana, the leaf beetles Exema dispar and Ophraella notata, and the gall gnat Dasyneura purpurea. Joe-Pye Weed species (Eutrochium spp.) are used by vertebrate animals only to a limited extent. The Swamp Sparrow and probably other birds consume the seeds, while hoofed mammalian herbivores (e.g., deer & cattle) eat the bitter-tasting leaves only when little else is available. Photographic Location: A wet sand prairie along a railroad at the Indiana Dunes National Lakeshore in NW Indiana. Most of the flowerheads of the photographed plant are still in the bud stage.
Comments: This wildflower is an impressive sight as it towers above the surrounding vegetation in open areas. Two other species in this genus occur in Illinois: Eutrochium purpureum (Sweet Joe-Pye Weed) and Eutrochium maculatum (Spotted Joe-Pye Weed). Like Hollow Joe-Pye Weed, these are tall plants with similar flowerheads and whorled leaves. Hollow Joe-Pye Weed can be distinguished from these species by the greater number of leaves per whorl (sometimes exceeding 5), the strong white bloom of its central stem, and the largely hollow interior of its central stem. The other two species of Joe-Pye Weed have at most 5 leaves per whorl, their central stems are either glabrous or hairy, but not strongly glaucous, and the interior of their central stems is only slightly hollow or filled with pith. Other scientific names of Hollow Joe-Pye Weed include Eupatorium fistulosum and Eupatoriadelphus fistulosus.
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Miss Chen
2018年06月18日
Miss Chen
Description: This herbaceous perennial plant is 1-4' tall, branching occasionally. The stems are square or round, and usually pubescent where new growth occurs, becoming glabrous with age. The hairless leaves are opposite or alternate along the stems, sessile or with short petioles, and up to 4" long and ½" across. They are narrowly lanceolate in shape, and sometimes have purple veins or spots. There are 4-8 teeth per centimeter along the margin of each leaf. From the upper axils of some of the upper leaves, there appears a single flower with a long calyx tube on a short stalk (peduncle). This calyx tube is pubescent and more or less erect, terminating in 4 lanceolate sepals that often tinted purple or pink. These sepals are a little shorter than the petals. The 4 notched petals are white or light pink, and span about 1/3" across. At the throat of the flower, there is a prominent pistil that is often knobby at the end, which is surrounded by several stamens. These flowers bloom for about a month during late summer or early fall. The calyx tube matures into an elongated seed capsule, which splits open to release a multitude of tiny seeds with small tufts of reddish brown hair. These seeds are distributed by the wind. The root system is fibrous and produces rhizomes, which enables this plant to spread vegetatively. Cultivation: The preference is full sun to light shade, and wet to moist conditions. The soil should contain lots of organic material to retain moisture. The foliage is rather fragile and can become damaged easily. This plant tolerates occasional flooding, and prefers rather cool conditions. Range & Habitat: The native Cinnamon Willow-Herb occurs occasionally in northern and central Illinois, but it is uncommon or absent in the southern section of the state. Its range extends further to the south than other Epilobium spp., which are usually restricted to northern Illinois. Habitats include poorly drained areas of black soil prairies, moist woodlands, woodland borders, and various kinds of wetlands, including marshes, bogs, fens, seeps, and edges of ponds, rivers, or drainage ditches. The Willow Herbs are pioneer species that thrive on some kind of disturbance, such as fire. Faunal Associations: The flowers probably attract bees and flower flies, which seek nectar and pollen. The caterpillars of various moths eat the foliage of Willow-Herbs, including Hyles lineata (White-Lined Sphinx), Eudryas unio (Pearly Wood Nymph), Anticlea multiferata (Many-Lined Carpet), and Scythris magnabella (Scythridid Moth sp.). The seeds are too small to be of much interest to birds. The foliage is non-toxic and occasionally eaten by mammalian herbivores, but it has low food value. Photographic Location: A poorly drained area of prairie in Meadowbrook Park, Urbana, Illinois. Comments: The flowers are rather small-sized, but they are often produced in abundance on large plants. The most distinctive characteristic of Cinnamon Willow-Herb is the reddish brown coloration of the tufts of hair on the seeds, which is responsible for the common name of this plant. Other Epilobium spp. have tufts of hair that are white or faded brown or grey. Cinnamon Willow-Herb also has serrated leaves with 4-8 teeth per centimeter, while other Willow-Herbs have leaves with either smooth edges or fewer teeth per centimeter. An unusual characteristic of the Willow-Herbs is the long calyx-tube of the flowers, which occurs behind the petals, and eventually becomes an elongated seed capsule. It looks like a flowering stalk (peduncle), but it is actually part of the flower where the ovaries are contained. In some members of the Mustard family and miscellaneous other plants, an elongated seed capsule develops in front of the petals. This is one way to determine whether the plant in front of you is a Willow-Herb, or a quite different species of plant.
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Miss Chen
2018年06月18日
Miss Chen
Description: This perennial wildflower is 2-5' tall and usually unbranched, except where the inflorescence occurs. There are no basal leaves. The terete central stem is light green, purplish red, or yellowish brown; the typical variety is glabrous or sparsely short-pubescent, while var. pubens tends to be more pubescent. Alternate leaves occur along the entire length of the stem; they are ascending and slightly recurved. The lowermost leaves are small and scale-like, while the remaining leaves are 3-5" long, ½-1" across, and relatively uniform in size as they ascend the stem. The leaves are elliptic or lanceolate-elliptic in shape, smooth and short-ciliate along their margins, and either sessile or with short petioles. The upper leaf surface is medium to dark green, while the lower surface is pale green or whitish green. The upper and lower leaf surfaces of the typical variety are glabrous to sparsely short-pubescent. The upper leaf surface of var. pubens is sparsely to moderately short-pubescent or canescent, while the lower surface is moderately to densely short-pubescent or canescent. The central stem terminates in a flat-headed panicle (compound corymb) of flowerheads about 3-12" across. Individual flowerheads are about ½" across, consisting of 5-12 ray florets that surround 12-25 disk florets. The petal-like corollas of the ray florets are white; the tubular corollas of the disk florets are yellow while in bloom, but become cream-colored or dingy white thereafter. Each disk floret has 5 spreading lobes. At the base of each flowerhead, there are 2-4 series of appressed floral bracts (phyllaries) that are narrowly oblong, green, and glabrous to short-pubescent. The branches of the inflorescence are light green or yellowish brown; they are either glabrous or short-pubescent. Leafy bracts about ½-1½" long occur along these branches; they are lanceolate to narrowly elliptic. The blooming period occurs from late summer into the fall, lasting 1-2 months. Both ray and disk florets are replaced by achenes with whitish tufts of hair. In each tuft of hair, the outermost hairs are bristly and short (less than 1 mm. in length), while the inner hairs are longer (3-6 mm. in length). Individual achenes are about 3 mm. (1/8") long, bullet-shaped, and sparsely short-pubescent. They are distributed by the wind. The root system is fibrous and rhizomatous. Occasionally, small colonies are formed from vegetative offsets. Cultivation: The preference is full or partial sun, slightly wet to moist conditions, and calcareous soil that contains sandy-loam. The pH should be slightly acidic. This wildflower can adapt to other kinds of soil, although they are not preferred. Range & Habitat: The native Flat-Topped Aster is occasional in NE Illinois and parts of central Illinois, while in the rest of the state it is uncommon or absent. Habitats include wet to moist sand prairies, moist sandy thickets, soggy meadows and openings in wooded areas, interdunal sloughs and swales near Lake Michigan, fens, and seeps. This wildflower is found in higher quality wetlands that are often sandy and calcareous. Faunal Associations: The nectar and pollen of the flowerheads attract long-tongued bees, short-tongued bees, wasps, flies, butterflies, beetles, and other insects. The foliage, plant juices, roots, and other parts of Flat-Topped Aster and other asters are eaten by many insects. The caterpillars of an uncommon butterfly, Chlosyne harrisii (Harris' Checkerspot), feed on Flat-Topped Aster specifically. Other insect feeders include caterpillars of the moths Acrocercops astericola, Astrotischeria astericola, and Carmenta corni (Aster Borer Moth); see the Moth Table for other moth species that feed on asters. Asters are also host plants of Macrosteles quadrilineatus (Aster Leafhopper), various aphids (primarily Uroleucon spp.), the plant bug Plagiognathus cuneatus, the lace bugs Corythucha marmorata and Galeatus spinifrons, the leaf beetles Microrhopala excavata and Exema canadensis, caterpillars of the butterflies Chlosyne nycteis (Silvery Checkerspot) and Phyciodes tharos (Pearl Crescent), larvae of Calycomyza humeralis (Aster Leafminer Fly), and larvae of small flies in the Tephritidae (Paroxyna albiceps, Tomoplagia obliqua, & Trupanea actinobola). Some vertebrate animals also feed on asters occasionally. The Ruffed Grouse and Wild Turkey feed on the seeds and foliage, while such songbirds as the Swamp Sparrow and Eastern Goldfinch also eat the seeds. The White-Tailed Deer and Cottontail Rabbit browse on the foliage; cattle, sheep, and other domesticated farm animals also browse on the foliage. Photographic Location: A wet sand prairie at the Indiana Dunes National Lakeshore in NW Indiana.
Comments: Flat-Topped Aster is an attractive wildflower that favors relatively open areas that are damp and sandy. Because of its flat-topped inflorescence, it is relatively easy to distinguish from other asters (Aster spp.) within the state. In particular, the prominent disk florets of the flowerheads are somewhat unusual in that they become dingy white or cream-colored shortly after they bloom, rather than orange-red or purple. The reason Flat-Topped Aster has been assigned to the Doellingeria genus is related to the different hair lengths of its tufted achenes: the outer hairs are less than 1 mm. in length, while the inner hairs are 3-6 mm. in length. Usually, asters have a uniform length for the hairs of their tufted achenes. An older scientific name of Flat-Topped Aster is Aster umbellatus.
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Miss Chen
2018年06月17日
Miss Chen
Description: This perennial wildflower forms arching stems about 3-8' long that are unbranched or sparingly so. The base of each stem is spongy and swollen if it is submerged in water, otherwise it is more constricted and angular. Each stem is pale green to red and either glabrous or pubescent; it tends to be slightly woody at the base, but dies down to the ground each winter (at least in the Midwest). Both opposite leaves and whorls of 3 leaves can occur along the stems; they are up to 6" long and 1½" across. The leaves are elliptic in shape and smooth along their margins. The upper leaf surface is medium to dark green and glabrous, while the lower surface is pale to medium green and either glabrous or pubescent. The leaves taper gradually into short petioles (¼" or less) and long narrow tips. Clusters of non-terminal flowers occur in the axils of the leaves. Individual flowers are up to 1" long and 1" across, consisting of 5 wrinkled purple petals, a short tubular calyx with 5 primary teeth alternating with 5 secondary teeth, 10 stamens of varying lengths, and a pistil with a style. Sometimes there are more calyx teeth and fewer stamens. The bell-shaped calyx is light green to cream-colored with rose tints; its primary teeth are ovate, while its secondary teeth are elevated above the primary teeth and they are linear. The slender secondary teeth are often contorted or crooked and they are ciliate along their margins. The pedicels of the flowers are light green, glabrous or pubescent, and short (about ¼" in length). There are 2 or more leafy bracts underneath each cluster of flowers; they are up to ¾" in lengthDistribution Map and lanceolate to ovate in shape. The blooming period occurs from mid-summer to early fall, lasting about 1-2 months. Each flower typically lasts only 1-2 days. During the autumn, the flowers are replaced by globoid seed capsules about ¼" across. Each capsule contains many chunky seeds; the seeds probably float on water. This wildflower reproduces clonally whenever its stem tips touch moist ground, where new plants will take root. As a result, colonies of clonal plants often form. Cultivation: The preference is full or partial sun, wet to moist conditions, and soil that is mucky, peaty, or sandy (or some combination of the preceding). Swamp Loosestrife is usually an emergent aquatic plant in stagnant or slow-moving water. However, it also colonizes damp ground near shorelines. Range & Habitat: Swamp Loosestrife is an uncommon wildflower that is found in scattered areas of Illinois. It is apparently absent in the NW and east-central areas of the state. Habitats include marshes and sandy marshes, swamps and sandy swamps, shorelines along ponds and small lakes, calcareous fens, and peaty bogs. Sometimes Swamp Loosestrife occurs on floating mats of vegetation in fens and bogs. It is usually found in high quality wetlands. Faunal Associations: The flowers are cross-pollinated by honeybees, bumblebees, Swallowtail butterflies, and probably other insects. These visitors obtain primarily nectar from the flowers. The caterpillars of Darapsa versicolor (Hydrangea Sphinx) and Eudryas unio (Pearly Wood Nymph) feed on the foliage, while the caterpillars of Papaipema cataphracta (Burdock Borer Moth) and Papaipema sulphurata (Decodon Borer Moth) bore through the stems. The range of the rare Decodon Borer Moth is restricted to northeastern United States. The seed capsules of Swamp Loosestrife are eaten by several ducks: the Mallard, Black Duck, Blue-Winged Teal, Green-Winged Teal, and Wood Duck. Muskrats like to feed on the swollen and spongy underwater stems.
Photographic Location: Shoreline of a small lake at the Indiana Dunes National Lakeshore in NW Indiana. Comments: Swamp Loosestrife is both large in size and attractive, especially when it is in bloom. Its two closest relatives in Illinois, the native Lythrum alatum (Winged Loosestrife) and introduced Lythrum salicaria (Purple Loosestrife), prefer somewhat drier areas of wetlands. Compared to Swamp Loosestrife, both of these species have very similar purple flowers, but they are both erect in their habit of growth, rather than arching.
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Miss Chen
2018年06月17日
Miss Chen
Description: This plant is a small much-branched shrub about 1½–3' tall. The bark of the woody branches is reddish brown to grayish brown, becoming shredded and ragged-looking with age. The alternate leaves are about ½–1" long and across; they are divided into 3-7 narrow pinnate lobes and their margins are smooth and revolute (folded downward). The lobes often appear to be palmate because they are crowded together; they are linear-oblong in shape. The upper surface of each leaf is medium to dark green and sparsely covered with fine silky hairs; the lower surface is pale green and more densely covered with fine hairs. The leaves have short petioles. The upper branches produce either individual or small clusters of yellow flowers. Each flower spans about ¾–1½" across; it has 5 yellow petals that are well-rounded, 5 yellowish green sepals, 15-20 stamens with flat golden anthers, and multiple carpels (female reproductive organs) that are crowded together in the center. The sepals are ovate in shape and sparsely covered with fine silky hairs. On a particular shrub, many flowers may bloom at once, or only a few flowers at a time. The blooming period occurs during the summer and early fall; a small colony of shrubs may remain in bloom for 2-3 months. Each flower is replaced by a capsule that contains several hairy seeds. Mature capsules become dark brown and persist into the winter. The root system consists of a woody taproot. Cultivation: Shrubby Cinquefoil prefers full sun, moist conditions, and a sandy or rocky soil containing calcium. The pH of the soil should be slightly acid to alkaline. Shrubby Cinquefoil can be cultivated in a garden soil containing loam or clay-loam, and it will adapt to drier situations.
Range & Habitat: Shrubby Cinquefoil is a rare plant in Illinois; it is found primarily in a few counties in the NE section of the state (see Distribution Map), where it is native. Habitats include fens, calcareous seeps, the upper portion of beaches and low sandy areas along Lake Michigan, sandy banks of interdunal ponds, moist dolomite prairies, and hill prairies. The last habitat is highly atypical. Shrubby Cinquefoil is typically found in calcareous wetlands near Lake Michigan and other Great Lakes; it rarely naturalizes in disturbed areas. Because of its attractive flowers and easy culture, Shrubby Cinquefoil is often cultivated as an ornamental plant in gardens and lawns. This circumboreal shrub is also native to Eurasia. Faunal Associations: According to observations of Müller (1873/1883) in Germany, the nectar and pollen of the flowers attract honeybees, Halictid bees, Sphecid wasps, and various flies. Observed fly visitors include: Soldier flies, Syrphid flies, Thick-headed flies, Flesh flies (Sarcophaga spp.), Blow flies (Lucilia spp.), Dung flies (Scatophaga spp., Sepsis spp.), and Anthomyiid flies. The caterpillars of the butterfly Lycaena dorcas (Dorcas Copper) feed on the foliage of Shrubby Cinquefoil, while the caterpillars of the moth Scopula limboundata (Large Lace Border) eat the flowers. Another insect that feeds on Shrubby Cinquefoil is the flea beetle Macrohaltica caurina. With the exception of goats, hoofed mammalian herbivores don't browse on this shrub to any significant extent.
Photographic Location: A roadside garden in Champaign, Illinois. Comments: Other scientific names for this species includes Potentilla fruticosa and Pentaphylloides floribunda. While the floral structure of Shrubby Cinquefoil (Dasiphora fruticosa) resembles that of the various cinquefoil species (Potentilla spp.), the latter are non-woody plants with smaller flowers (¼–¾" across). Another species, Three-toothed Cinquefoil (Sibbaldiopsis tridentata), is a low woody plant with small white flowers; it more closely resembles a wild strawberry (Fragaria sp.) than Shrubby Cinquefoil.
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Miss Chen
2018年06月17日
Miss Chen
Description: This herbaceous vine is a summer annual up to several feet long that branches occasionally to abundantly. It sprawls across other vegetation that it parasitizes, extracting water and nutrients through clinging suckers (haustoria). Sometimes this vine is so robust that it forms a dense maze of stems that partially obscures the vegetation underneath. The stems are orange, terete, glabrous, and occasionally warty. The alternate leaves are reduced to minute scales or they are absent. Small clusters of flowers develop at intervals along the stems on greenish yellow peduncles and pedicels that are glabrous. The pedicels of the flowers are very short (less than 1 mm. in length). Each flower spans about 3 mm. across, consisting of a short-tubular calyx with 4 obtuse lobes, a short-tubular corolla with 4 ascending to erect lobes, 4 stamens, and a pistil with a pair of divergent styles. Less commonly, a flower may have 5 calyx lobes, 5 corolla lobes, and 5 stamens. The calyx is light green or pale yellow and glabrous, while the corolla is white and its lobes are ovate with blunt tips. The lobes of the calyx extend to about the sinuses of the corolla or a little less. Inside the corolla at the base of the stamens, there are fringed floral scales (requires at least a 10x hand lens to see). The blooming period occurs from mid-summer to early fall, lasting about 1½ months. The flowers may be slightly fragrant, and they are self-fertile. The flowers are replaced by globoid seed capsules spanning 3-5 mm. across; they are surrounded by their persistent calyces and the withered remnants of their corollas. Immature seed capsules are light green or pale yellowish green, but they become brown at maturity. Each capsule has 2 cells and contains up to 2 seeds per cell. The seeds are 1.0-1.5 mm. in length and ovoid in shape. While a young seedling has a rudimentary root system, it soon withers away after the seedling attaches itself to a suitable host plant. Without a host plant, the seedling soon dies as it lacks chlorophyll.Stems & Buds
Cultivation: This parasitic vine is typically found in moist to wet open areas where the soil contains abundant organic material and possibly some silt or sand. Favored host plants are smartweeds (Persicaria spp.), but many other species of plants are also parasitized. During certain times of the year, the host plants may be found in shallow standing water. Smartweed Dodder can spread aggressively and seriously weaken its host plants. The small seeds of dodders (Cuscuta spp.) can remain viable for 5-7 years (Georgia, 1913). Range & Habitat: The native Smartweed Dodder is scattered across most areas of Illinois, except possibly the NW section of the state (see Distribution Map). In those areas where this species occurs, it is uncommon to occasional. Habitats include moist to wet prairies, soggy thickets along rivers, fens, sandy marshes, and other wet places. Faunal Associations: The floral nectar of dodders (Cuscuta spp.) attracts Halictid bees (Lasioglossum spp.), plasterer bees (Colletes spp.), and other small bees (Robertson, 1929). According to Müller (1873/1883), the flowers can attract moths. According to Georgia (1913), the seeds of these parasitic vines can pass through the digestive tracts of grazing animals (e.g., cattle, horses, etc.) and remain viable. Thus, these animals may help to spread these vines into new areas. Photographic Location: Cowle's Bog (actually, a fen) at the Indiana Dunes National Lakeshore in NW Indiana. Comments: Because of its bright orange stems, this is a conspicuous vine, especially when it develops into a sprawling mass that resembles a large pile of abandoned tangled string. This is quite a sight to behold. Smartweed Dodder can be distinguished from most species of dodders (Cuscuta spp.) by its 4-parted flowers (4 calyx lobes, 4 corolla lobes, and 4 stamens). Two exceptions are Hazel Dodder (Cuscuta coryli) and Buttonbush Dodder (Cuscuta cephalanthi), which also have 4-parted flowers. These latter two species have yellow stems, rather than bright orange stems. Compared to Smartweed Dodder, the corolla lobes of Hazel Dodder are more erect with acute and incurved tips, while the corolla lobes of Buttonbush Dodder are widely spreading. Aside from these species, other dodders in Illinois have 5-parted flowers (5 calyx lobes, 5 corolla lobes, and 5 stamens).
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Miss Chen
2018年06月17日
Miss Chen
Buttonbush Dodder on a Host PlantDescription: This herbaceous parasitic vine is a summer annual that is several feet along. It clings to adjacent vegetation using suckers (haustoria) on its stems. Lacking chlorophyll and a significant root system, this vine is dependent on its host plants for water and nutrients. The stems are usually pale yellow, sometimes becoming pale orange with age; they are glabrous and terete. Alternate leaves along the stems are reduced to tiny scales, or they are absent altogether. Small clusters of nearly sessile flowers occur at intervals along the stems. The peduncles and pedicels of these flowers are pale yellow to pale orange and glabrous. Each flower spans about 3 mm. (1/8" across), consisting of a white corolla with 4 lobes, a light green calyx with 4 lobes, 4 stamens, and an ovary with a pair of divergent styles. Less often, a flower may have a corolla with 5 lobes, a calyx with 5 lobes, and 5 stamens. The lobes of the corolla are deltate-ovate in shape and ascending to widely spreading. The calyx is glabrous with rounded lobes; it is shorter than the corolla. Both the corolla and calyx are short-tubular and bell-shaped (campanulate). At the base of the stamens within the corolla, are fringed scales that require at least a 10x hand lens to see. There are no floral bracts underneath the flowers. The blooming period occurs from mid-summer to early fall, lasting about 1½ months. There may be a mild floral fragrance. In the absence of insect pollinators, the flowers are self-fertile; they are replaced by globoid seed capsules spanning 3-5 mm. across. The glabrous seed capsules are surrounded by the withered remains of their corollas; they are initially light green, but turn brown at maturity. Each capsule is 2-celled, and each cell contains 2 seeds. The capsules eventually split open to release their seeds. The seeds are about 1.5 mm. long (or slightly more), ovoid in shape, and slightly compressed. There is a rudimentary root system at the seedling stage, but this is abandoned shortly after a suitable host plant has been found. If no host plant is found, the seedling dies within a few days. Cultivation: This vine is typically found in wet to moist areas that are exposed to sun or partial sun. It cannot survive without a suitable host plant, although apparently many species of moisture-loving plants and shrubs can serve this purpose. Examples of suitable host plants include buttonbush (Cephalanthus occidentalis), water willow (Justicia americana), false nettle (Boehmeria cylindrica), asters (Aster spp.), goldenrods (Solidago spp.), bugleweed (Lycopus spp.), and even horsetails (Equisetum spp.). Soil type depends on the preferences of host plants, but can consist of sand, silt, loam, or muck. This vine can significantly weaken any host plant that it comes in contact with.
Range & Habitat: The native Buttonbush Dodder is scattered throughout Illinois, where it is uncommon to occasional (see Distribution Map). Habitats include floodplain woodlands, swamps, soggy thickets along rivers, marshes, and wet prairies. This vine occurs in both sandy and non-sandy wetlands. Faunal Associations: Robertson (1929) observed a small Halictid bee, Lasioglossum coriaceus, sucking nectar from the flowers of Buttonbush Dodder. Other small bees also visit the flowers for nectar, and possibly some moths (Müller, 1873/1883). According to Georgia (1913), the seeds of dodders can pass through the digestive tracts of grazing animals (e.g., cattle & horses) and remain viable. Thus, these animals may help to distribute these vines into new areas.
Photographic Location: A sandy marsh at the Heron Boardwalk in Vermilion County, Illinois. Comments: Buttonbush Dodder has a fairly typical appearance for the parasitic vines of this genus, although it is somewhat unusual in having 4-parted flowers (4-lobed corollas, 4-lobed calyces, and 4 stamens), rather than 5-parted flowers (5-lobed corollas, 5-lobed calyces, 5 stamens). Unlike other dodder species (Cuscuta spp.) with 4-parted flowers, the corolla lobes of Buttonbush Dodder usually spread outward rather than remain erect. Thus, it is a fairly easy species to identify during the blooming period. Some taxonomists have placed dodder species in the Morning Glory family (Convolvulaceae).
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Miss Chen
2018年06月17日
Miss Chen
Description: This perennial plant is 1-2½' tall, branching occasionally to abundantly. The stems are light green, terete, and appressed-pubescent. The opposite leaves are up to 3" long and 2" across; they have short slender petioles. The leaf blades are oval-cordate, oval, or ovate in shape, while their margins are dentate or dentate-crenate. The upper blade surface is light green and glabrous with a conspicuous network of veins. The upper stems terminate in flat-topped clusters of flowerheads. Each cluster of flowerheads spans about 1-3" across. Each flowerhead has about 40-50 disk florets that are pink, lavender, or blue. Each floret has a tiny tubular corolla with 5 spreading lobes and a strongly exerted style that is divided into two filiform parts. Around the base of each flowerhead, there are several floral bracts (phyllaries) that are arranged in 1-2 series; they are light green and linear in shape. The branches underneath each flowerhead cluster are light green and terete. The blooming period occurs from mid-summer to early autumn, lasting about 1-2 months. There is no noticeable floral scent. Afterwards, the florets are replaced by achenes with small tufts of hair; they are distributed by the wind. The root system is highly rhizomatous; this plant readily forms colonies. Cultivation: The preference is full sun to light shade, moist conditions, and soil containing loam or silt. There should be sufficient organic material in the soil to retain moisture. This plant can spread aggressively in moist open ground, otherwise it presents few problems. Drought tolerance is poor. Propagation is by seed or division of the rhizomes.
Range & Distribution: The native Mistflower occurs in the southern half of Illinois and the Chicago area (see Distribution Map). It is fairly common in southern Illinois, but uncommon or absent elsewhere. This plant was introduced into the Chicago area. Some local populations in the wild are probably the result of seeds or plants that have escaped cultivation. Habitats include river-bottom prairies, moist open woodlands, gravelly seeps, borders of lakes and rivers, moist meadows in wooded areas, bases of bluffs, and ditches. This plant usually occurs in poorly drained areas and near sources of water. Faunal Associations: The flowers attract long-tongued bees, butterflies, and skippers. Other occasional visitors include short-tongued bees, various flies, moths, and beetles. These insects seek nectar primarily, although the bees often collect pollen. Insects that feed on Eupatorium spp. (Bonesets) may also feed on Mistflower. Insect feeders of this group of plants include the caterpillars of such moths as Haploa clymene (Clymene Moth; eats foliage), Phragmatobia lineata (Lined Ruby Tiger Moth; eats foliage), Carmenta bassiformis (Eupatorium Borer Moth; bores into roots), and Schinia trifascia (Three-Lined Flower Moth; eats florets & developing seeds). Mammalian herbivores rarely consume Mistflower because of its bitter foliage.
Photographic Location: The photograph was taken along a drainage canal at Kaufman Lake Park in Champaign, Illinois. Comments: Mistflower has attractive delicate flowers that are colored in pastel shades of pink, lavender, or blue. For this reason, it is often grown in flower gardens. This plant is closely related to the white-flowered Bonesets (Eupatorium spp.), and sometimes it is still referred to as Eupatorium coelestinum. Mistflower can be distinguished from the Bonesets primarily by its colorful flowers, relatively short stature, and broad opposite leaves that are heavily veined. While species of Joe-Pye Weed (Eupatoriadelphus spp.) have similar colorful flowers, they are taller plants with whorled leaves. All of these species are similar in that their flowerheads consist entirely of disk florets.
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Miss Chen
2018年06月16日
Miss Chen
Description: This perennial wildflower is 1½–6' tall, branching occasionally along the upper half of its length. The rather stout stems are light green to reddish purple (often the latter), terete to slightly grooved, and evenly covered with stiff spreading hairs. The alternate leaves are up to 6" long and 1¾" across, becoming gradually smaller along the upper half of each plant; they are narrowly lanceolate to lanceolate with poorly defined remote teeth along their margins. The leaves are yellowish green, medium green, or purple (sometimes the latter color during the fall); they are usually glabrous, except for some hairs along the central veins of their lower sides. Most leaves clasp the stems, although some of the smaller upper leaves are sessile. The central stem terminates in a panicle of flowerheads; some lateral stems may produce smaller panicles of flowerheads. The branches of each panicle are ascending and usually hairy. Along these branches, there are linear-lanceolate leafy bracts up to 1" long. The outer branches terminate in flowerheads about ¾–1¼" across, consisting of 30-50 ray florets and a similar number of central disk florets. The petal-like rays are usually lavender, pale blue-violet, or purple (less often white); they are widely spreading and very slender. The tubular disk florets are 5-lobed; they are initially yellow, but later become dull red. At the base of each flowerhead, there are several overlapping bracts that are linear in shape, green, and hairless; they are rather loosely assembled around the base of the flowerhead and slightly spreading. The blooming period occurs from late summer into the fall and lasts about 2 months. Both disk and ray florets are fertile. The florets are replaced by bullet-shaped achenes about 1.5 mm. long that have small tufts of white hair; they are distributed by the wind. The root system is fibrous and short-rhizomatous, sometimes forming a small caudex on older plants. Cultivation: The preference is full or partial sun and wet to moist conditions. The soil should contain some organic material to retain moisture and it should be reasonably fertile. Sometimes, the leaves become diseased and rather battered in appearance by the end of the year; the lower leaves may turn brown and fall off in response to droughty conditions. The size of individual plants can be highly variable. Range & Habitat: The native Swamp Aster is occasional in the northern half of Illinois, while in the southern half of the state it is uncommon or absent. Habitats include soggy thickets along streams, open swamps, fens and calcareous seeps, sedge meadows, and other wetlands. Swamp Aster is often found in higher quality wetlands where the native flora is still intact. It is primarily a boreal species that is found around the Great Lakes and other cool areas. Faunal Associations: The nectar and pollen of the flowerheads attract a wide variety of insects, including honeybees, bumblebees, other miscellaneous bees, various wasps, bee flies and other miscellaneous flies, and various butterflies, skippers, and moths. The oligolectic bees, Andrena asteris and Andrena hirticincta, suck nectar and collect pollen from the flowerheads of Swamp Aster. Other insects feed on the foliage, suck plant juices, bore through the stalks and roots, or gnaw on the flowers and developing seeds of Symphyotrichum spp. (asters). These species include Microrhopala xerene and other leaf beetles, several aphids (mostly Uroleucon spp.), the stinkbug Trichopepla semivittata, the leafhopper Macrosteles quadrilineatus, the plant bug Plagiognathus cuneatus, Poecilocapsus lineatus (Four-Lined Plant Bug), Lygus lineolaris (Tarnished Plant Bug), and the larvae of Calycomyza humeralis (Aster Leafminer Fly). In addition to these insects, a large number of moth caterpillars feed on asters (see the Moth Table for a listing of these species), as do the caterpillars of the butterflies Chlosyne nycteis (Silvery Checkerspot) and Phyciodes tharos (Pearl Crescent). Among vertebrate animals, the Wild Turkey eats the seeds and leaves occasionally, while the White-Tailed Deer and Cottontail Rabbit browse on the foliage. Photographic Location: A low area along a pond at Weaver Park in Urbana, Illinois.
Comments: Except for Symphyotrichum novae-angliae (New England Aster), Swamp Aster has larger flowerheads than other Symphyotrichum spp. (asters) in Illinois and its flowerheads usually have more rays (about 30-50). It is a pretty hefty plant that is about as tall as New England Aster. This latter species has glandular hairs along its stems and its leaves are less shiny than those of Swamp Aster. Swamp Aster is divided into two varieties or subspecies: the typical variety/subspecies has evenly hairy purple stems, while variety/subspecies firmus (or lucidulous) has light green stems that are either hairless or the hairs are arranged in lines; sometimes the color of the stems is not reliable in making this distinction. The rhizomes of the typical variety/subspecies are supposed to be short, while the rhizomes of variety/subspecies firmus (or lucidulous) are supposed to be long. Some authors (e.g., Mohlenbrock, 2014), divide Swamp Aster into two separate species: Symphyotrichum puniceum and Symphyotrichum firmum. They have a similar distribution and are about equally common in Illinois. The preceding descriptive information and photos apply to Symphyotrichum puniceum puniceum. A scientific synonym of Swamp Aster is Aster puniceus.
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